Genus Pseudoceros
Phylum Platyhelminthes:
Flatworms
After reviewing this section, the
student should be able to discuss the physiology of flatworms,
and distinguish among the classes
Turbellaria,
Trematoda, and Cestoda.
There
are about 20,000 species of Platyhelminthes, most of them parasitic. All
flatworms are acoelomate,
triploblastic and bilaterally symmetrical.
The flatworms
are the simplest animals with bilateral symmetry and three distinct germ
layers, yet their tissues are specialized for various functions, and two
or more types of cell tissues may combine to form organs. Unlike
most other bilaterally symmetrical animals, the flatworms have a digestive
cavity with only one opening. Since the animals cannot feed, digest,
and eliminate undigested residues simultaneously, food cannot be processed
continually.
Flatworms have
solid bodies, with no circulatory system for the transport of oxygen and
food molecules, so all cells must be within diffusion distance of sources
of oxygen and food. Their body structure is well suited to fulfilling
these requirements. The body is flattened, which keeps the cells
close to the external oxygen supply. Also, the digestive cavity is
branched, carrying food particles to all regions of the body.
About 13,000
species of flatworms have been described, and they are placed in three
classes: class Turbellaria,
which contains mostly free-living (nonparasitic) forms; class Trematoda,
encompassing parasitic flukes; and class Cestoda,
with tapeworms (also parasitic).
Class Turbellaria
The
free-living
(non-parasitic) flatworms form a large and varied group. Turbellarians
are predominately free-living and aquatic. They have an incomplete digestive
tract, in which the mouth leads to a pharynx, then to temporary spaces
containing cells that take in food particles by phagocytosis. Digestion
is intracellular. These animals move by laying down slime from special
skin glands, then gliding along these "slime trails" by the beating of
epidermal cilia.
Turbellarians are mostly carnivorous, preying on tiny invertebrates that they locate by means of their chemoreceptors. They are a diverse group, including over 4500 known species divided into twelve orders.
Class Trematoda
All
trematodes are parasitic,
and most adult trematodes parasitize vertebrates. Around 9000 species have
been described. Their body is covered with a tegument, a peculiar kind
of epidermal arrangement in which the main cell bodies are deep, separated
from the cytoplasm that lies next to the exterior by a layer of muscle
(but connected to the exterior layer by cellular processes. The tegument
lacks cilia in adults. Trematodes are characterized by one or two
suckers or hooks on their anterior ends by which they fasten onto their
victims. They are like turbellarians in having a relatively well developed
alimentary canal, and their muscular, excretory, and reproductive systems
are also relatively complete.
Most
trematodes have complex life cycles, with larval stages parasitizing one
or more species that are different from host of adults. Most trematodes
are endoparasites (parasites which live inside the host's body). They include
several parasites that have an enormous impact on human populations, such
as human liver flukes and the blood flukes that cause schistosomiasis.
The symptoms of schistosomiasis are caused by the eggs, which have sharp
spines on the surface and are laid in the capillaries of the bladder wall
or intestines. The eggs lodge in the liver and spleen, blocking blood
vessels, and their sharp spines tear the surrounding tissues, causing hemorrhages.
Schistosomiasis now affects some 200 to 300 million people in 71 countries
in Asia, Africa, and South America.
Class Cestoda
The
cestodes, or tapeworms, differ in a number of ways from other flatworms.
Their bodies are long and flat, made up of many segments called proglottids.
Each proglottid is a reproductive unit, essentially a factory to produce
gametes. Cestodes' teguments are covered with tiny projections, microvilli,
which increase its surface area and thereby increase its ability to absorb
nutrients from a host. Digestive tracts are absent completely, and tapeworms
also lack mouths and digestive enzymes. At the tapeworm's anterior end
is a specialized segment called a scolex, which is usually covered with
hooks or suckers and serves to anchor it to the host. The tapeworms
merely hang on and absorb predigested food molecules through their skin.
Tapeworms
are found in the intestines of many vertebrates, including humans, and
may grow as long as five or six meters. They cause illness not only
by encroaching on the food supply but also by producing wastes and by obstructing
the intestinal tract. The most common human tapeworm, the beef tapeworm,
infects people who eat the undercooked flesh of cattle that have grazed
on land contaminated by human feces containing tapeworm segments.
All of the 5000
or so known species of tapeworms are endoparasites.
Most require at least two hosts, with the host of the adult tapeworm a
vertebrate. Intermediate hosts are often invertebrates. A number of tapeworm
species inhabit humans.
Thanks so much to these extremely
helpful sites for the information and/or pictures they provided!
Check them out for even more in depth information, pictures, and essays.
http://animaldiversity.ummz.umich.edu/platyhelminthes.html
http://www.sidwell.edu/us/science/vlb5/Labs/Classification_Lab/Eukarya/Animalia/Platyhelminthes/
http://biology.unm.edu/biology/bio404/Platyhelminthes.html